What are the inclusion relationships if any, between the classes Pspace, PLS, PPP, PPA and PPAD I have been thinking about Pspace in conjunction with searching for a Natural Notion of Stablilty for Complex Dynamical Systems.

A natural question in this direction is the Nash equilibrium. Surprizingly, it appears that Games in Normal Form have Nash equilibria which is PPAD complete. This has led myself to wonder about the inclusion relationships of various associated Complexity Classes. 

Thanks in advance.
Strictly speaking, PSPACE contains decision problems, while the others are search problems, so they live in different zoos, but FPSPACE would of course contain all the other problems. Then PPAD is contained in both PPA and PPP, whose relationship is unknown, but were separated by an oracle in Beame et al: Relative complexity of NP search problems. Here the PLS class was not considered, that was separated by an oracle later in Morioka: Relative complexity of local search heuristics and the iteration principle from all the other three classes. (For me this latter paper was too hard to read.) Without oracles, nothing is known.