What is this 运行 in ...国内的汽车行业与汽车市场运行环境发生了前所未有的改变? 中国加入世界贸易组织后，国内的汽车行业与汽车市场运行环境发生了前所未有的改变。


Sometimes in colloquial English we say 'shift' to mean 'sell'. Is '运行环境‘ something like that?

运行环境 maybe 'sales environment' or ‘turnover' (交易额） ??
国内的汽车行业与汽车市场运行环境发生了前所未有的改变 

This segment I would probably translate as "The domestic automobile industry and the operating environment of the automobile market underwent unprecedented changes." 

国内的 domestic
汽车行业 automobile industry
与 and
汽车市场运行环境 operating environment of the automobile market
发生了 happened/underwent
前所未有的 unprecedented
改变 changes

Hope that helps!