Improve error messages.
Improve error messages.

See issue: https://www.ebi.ac.uk/panda/jira/browse/GOA-2378

ebi-uniprot/QuickGOBE,ebi-uniprot/QuickGOBE,ebi-uniprot/QuickGOBE,ebi-uniprot/QuickGOBE,ebi-uniprot/QuickGOBE