all popsizes should be in the updown operator (when used)
all popsizes should be in the updown operator (when used)

Anaphory/beast2,tgvaughan/beast2,tgvaughan/beast2,CompEvol/beast2,CompEvol/beast2,CompEvol/beast2,Anaphory/beast2,Anaphory/beast2,Anaphory/beast2,tgvaughan/beast2,tgvaughan/beast2,CompEvol/beast2